Species Richness & Diversity (SL IB Environmental Systems & Societies (ESS))

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Alistair

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Alistair

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Biology & Environmental Systems and Societies

Species Richness & Diversity

What is species richness?

  • Species richness is the number of species in a community or defined area and can be a useful comparative measure in some cases
  • However, in other cases, species richness can be a misleading indicator of diversity as it does not take into account the number of individuals of each species
  • Once the abundance of each species in an area has been recorded, the results can be used to calculate the species diversity for that area
    • Species diversity looks at the number of different species in an area but also the evenness of abundance across the different species (i.e. their relative abundances)

Species Richness vs Diversity

  • An index of diversity is a measurement that describes the relationship between the number of species present and how each species contributes to the total number of organisms that are present in that community
    • It is a much more informative measurement than species richness and conservationists often favour the use of an index of diversity as it takes into account both species richness and evenness
    • A commonly used index of diversity is known as Simpson’s Diversity Index

Example

  • Area 1 and Area 2 both contain 4 tree species
  • However, Area 2 is actually dominated by one species and in fact, one of the species is very rare (only one individual)
  • Although the two areas have exactly the same species richness, Area 1 has a higher species evenness (and therefore a higher overall species diversity) than Area 2
  • This example illustrates the limitations of using just species richness on its own

Species richness and evenness
Area 1 and Area 2 have the same species richness but different species evenness. As it has a higher species evenness, the overall species diversity of Area 1 is higher than that of Area 2, as species diversity takes both richness and evenness into account

Calculating Simpson’s Diversity Index

  • A group of students used the kick sampling technique to collect, identify and count the invertebrates inhabiting a river
  • Samples were obtained from different sites along the course of the river
  • The data was used to calculate the Simpson's Diversity Index at two different river sites
  • This index of diversity is useful when comparing two similar habitats, or the same habitat over time
  • The formula for calculating Simpson's Diversity Index, D, is:

Species Mean number of organisms per m2 of river bed
Site A Site B
Mite 14 0
Snail 9 0
Leech 3 26
Worm 0 6
Flat worm 132 9
Mayfly nymph 43 0
Olive mayfly nymph 154 0
Midge Larva 0 10
Blackfly larva 77 0
Caddis larva 15 1
Fish 1 0
Freshwater shrimp 211 6
Water hog louse 0 40

Site A

Species Number (n)  n (n-1)
Mite 14 182
Snail 9 72
Leech 3 6
Worm 0 0
Flat worm 132 17 292
Mayfly nymph 43 1 806
Olive mayfly nymph 154 23 562
Midge Larva 0 0
Blackfly larva 77 5 852
Caddis larva 15 210
Fish 1 0
Freshwater shrimp 211 44 310
Water hog louse 0 0
Total N= ∑n= 659 ∑n(n-1)= 93 292

bold italic D equals bold italic space fraction numerator N stretchy left parenthesis N minus 1 stretchy right parenthesis over denominator capital sigma n stretchy left parenthesis n minus 1 stretchy right parenthesis end fraction bold italic equals fraction numerator space 659 stretchy left parenthesis 658 stretchy right parenthesis over denominator 93292 end fraction bold italic space equals space 4.65

Site B

Species Number (n)  n (n-1)
Mite 0 0
Snail 0 0
Leech 6 30
Worm 26 650
Flat worm 9 72
Mayfly nymph 0 0
Olive mayfly nymph 0 0
Midge Larva 10 90
Blackfly larva 0 0
Caddis larva 1 0
Fish 0 0
Freshwater shrimp 6 30
Water hog louse 40 1 560
Total N= ∑n= 98 ∑n(n-1)= 2 432



bold italic D equals bold italic space fraction numerator N stretchy left parenthesis N minus 1 stretchy right parenthesis over denominator capital sigma n stretchy left parenthesis n minus 1 stretchy right parenthesis end fraction bold italic equals fraction numerator space 98 stretchy left parenthesis 97 stretchy right parenthesis over denominator 2432 end fraction bold italic space equals space 3.91

  • Site A was located just 5 km downstream from the river’s source
  • Site B was located 50 m downstream from a sewage inlet pipe
  • The lower diversity index for site B reflects the stress placed upon the river as a consequence of the pollution load from the sewage inlet point; although the river is recovering from the pollution, sensitive species are unable to tolerate the unfavourable abiotic conditions and species diversity is reduced

Exam Tip

Remember, this index of diversity is only useful when comparing two similar habitats, or the same habitat over time.

You will be provided with the formula for Simpson’s Index in the exam but you need to know how to use it to calculate Simpson’s Diversity Index for example sets of data.

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Alistair

Author: Alistair

Alistair graduated from Oxford University with a degree in Biological Sciences. He has taught GCSE/IGCSE Biology, as well as Biology and Environmental Systems & Societies for the International Baccalaureate Diploma Programme. While teaching in Oxford, Alistair completed his MA Education as Head of Department for Environmental Systems & Societies. Alistair has continued to pursue his interests in ecology and environmental science, recently gaining an MSc in Wildlife Biology & Conservation with Edinburgh Napier University.